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Sausage Rules
(slrpnk.net)
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They were squeezed out
I thought so as well. Maybe the other side of the protruding sausages were progressively squeezed out by the dough starting from the tip, and it's probably due to the proximity to other rolls in the back, or maybe the center of the oven was much hotter and ended up cooking (and growing) the dough unevenly. That explains why the tip of the sausages looks popped.